What is the difference between the $ operator in Haskell and Purescript

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This document, about the differences between Haskell and Purescript, says the following about the $ operator

GHC provides a special typing rule for the $ operator, so that the following natural application to the rank-2 runST function is well-typed:

runST $ do
  ...

PureScript does not provide this rule, so it is necessary to either

omit the operator: runST do ... or use parentheses instead: runST (do ...)

Which typing rule is meant by this? And how does this impact the example of runST?

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