How to track down what is causing the errors on my site

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When i go to several pages on my site I now get this error. 90% of the time my site works and then 10% i get this error message. By just refreshing the page the content will eventually appear. The error appears completely random.

The problem I have is that I don't know how to diagnose the cause of the fault from here. How would I review the stack to work out what is causing this? What would be the best way to track down the cause of this fault?

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