Why does the closure for `take_while` take its argument by reference?

Here is an example from Rust by Example:

fn is_odd(n: u32) -> bool {
    n % 2 == 1
}

fn main() {
    println!("Find the sum of all the squared odd numbers under 1000");
    let upper = 1000;

    // Functional approach
    let sum_of_squared_odd_numbers: u32 =
        (0..).map(|n| n * n)             // All natural numbers squared
             .take_while(|&n| n < upper) // Below upper limit
             .filter(|n| is_odd(*n))     // That are odd
             .fold(0, |sum, i| sum + i); // Sum them
    println!("functional style: {}", sum_of_squared_odd_numbers);
}

Why does the closure for take_while take its argument by reference, while all the others take by value?