What does the period mean when used with ~ (in a formula)?

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From the FSelector manual:

data(iris)
subset <- cfs(Species~., iris)
f <- as.simple.formula(subset, "Species")
print(f)

Specifically, I mean the one in "Species~.".

Now, it's awfully tough to Google how a bit of punctuation is used (for me anyway) and I couldn't anything. This code is unclear.

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